Pollen: Spheroid, round, pantoaperturat, reticulate. Thick muri, narrow brochi. Aperturmembranes are heavily ornamented: operculat. The opercel are occupied with verrucate sculptures. Pores usually round, pore diameter ca. 8.3 (7.8-8.8) microns. Average pore spacing approximately 14.4 (11.5-19.1) microns. Diameter of the PK without the opercel.